Richard Holmes

|Research Professor
Academic Appointments
  • Research Professor of Biology

  • Ronald and Deborah Harris Professor of Environmental Biology Emeritus

[Please note: As I am now retired, I am no longer accepting new graduate students]

My research focuses on the ecology and behavior of migratory passerine birds in their north temperate breeding areas in New England and their neotropical wintering quarters in the Caribbean and Central America. Our current research examines climate effect on bird distribution, abundance, and demography and will contribute to our understanding of the factors affecting neotropical migrant bird populations, some of which are currently declining.


340 Life Sciences Center


Biological Sciences


  • B.S. Humboldt State University
  • Ph.D., University of California, Berkeley

Selected Publications

  • Holmes, R.T., and G. E. Likens. 2016. Hubbard Brook: The Story of a Forest Ecosystem. Yale University Press, New Haven. pp 1-270.

  • Townsend, A. K., E. G. Cooch, T. S. Sillett, N. L. Rodenhouse, R. T. Holmes, and M. S. Webster. 2016. Contrasting effects of warm springs and global climate cycles on population dynamics of a migratory songbird. Global Change Biology 22: 544-555 (doi: 10.1111/gcb.13053).

  • Marra, P. P., C. E. Studds, S. Wilson, T. S. Sillett, T. W. Sherry, and R. T. Holmes. 2015. Non-breeding season habitat quality mediates the strength of density-dependence for a migratory bird. 2015. Proceedings of the Royal Society London B 282,

  • Sherry, T.W., S. Wilson, S. Hunter, and R.T. Holmes. 2015. Impacts of nest predators and weather on reproductive success and population limitation in a long-distance migratory songbird. Journal of Avian Biology 46: 559- 569.

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